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RE: the mutation bug - it seemed as though cells would only ever attempt a single shared mutation across the entire species and the sim was failing to validate this against the editable conditions to determine viability. They each began to produce the same doomed mutation and the lineage would completely stagnate. Absolutely had me banging my head against the wall.
I hadn't actually made any initial changes to the default constants. Thankfully, the issue only occurs in the Vulkan runtime and so I saw immediate progress when switching to OpenGL, minus a few frames.