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The Horsemen of Apocalypse are a team of fictional supervillain characters that appear in comic books published by Marvel Comics. The group consists of four individuals (usually mutants) that have been genetically altered and mentally conditioned to serve the ancient mutant, Apocalypse, either willingly or forcibly. They are enhanced or endowed with new abilities, and are always given the same titles based upon the biblical Four Horsemen (Death, Famine, Pestilence (replacing the biblical Conquest), and War). While Apocalypse has empowered other individuals to do his bidding, the Four Horsemen remain his elite minions, always playing a key role in his plans.