This guide is intended to help players who might struggle with one of the colourfully-named bosses of Blasphemous 2. For the sake of consistency I decided to write strategies for all of them even if the strategy is obvious or the boss is easy to beat.

The Attacks section of each boss has the following structure:
Attack #
Move: What the attack does. (What the stronger version of the attack does)
Signalling: How can you tell the attack is coming.
Evade: How to avoid getting hit.

The attacks are numbered and referred in Strategy section as A1 for Attack 1 etc. Strategy section describes how to defeat the boss. Certain attacks are just stronger modifications of previous attacks, so I merged them into one attack; the stronger variant is described in parentheses.

To beat a boss, you usually have to damage it until you deplete its health bar which is visible in the lower part of the screen (stating the obvious here). Certain bosses have multiple phases and each one usually has its own full health bar. I tried to estimate each boss' total healthpoints using the damage numbers display so this estimate is also listed in each section.

One thing about Blasphemous 2 is that different players' playthroughs can differ quite a bit as a player might not have an upgrade, a prayer, a bead or a figure when reaching a boss, which makes writing the strategies slightly harder. I focused mainly on describing ways to avoid the enemy attacks regardless of the player's status, leaving the method of dealing damage to the boss for the readers to figure out, though occassionally I included tips. It is a good idea to search the available area for upgrades and items to help you to face the boss as strong as you can. You can equip rosary beads to improve your resistance to various types of damage, so pay attention to what kind of element each boss uses and change your equipment accordingly.

At the end of certain sections I added my personal thoughts about the bossfight.

There is a NPC named Yerma which you encounter early in the game and for certain bosses, you can request her help for the fights (this is similar to Blasphemous 1). I mention the availability of Yerma in the appropriate boss sections. I am not aware of any downside of asking her for help, though she usually shows up only in the beginnings of the fights.

Beating most bosses gives you Tears of Atonement and Marks of Martyrdom. I will try to add the amount in a future guide update.

For each boss you also get an achievement, but I don't list them in the guide.

Any feedback or improvement suggestions are welcome, both for content and for language (since English is not my first language).

Location: Repose of the Silent One
Healthpoints estimate: 600
Tears of Atonement: 500
Marks of Martyrdom: 1

Attacks
Attack 1
Move: The boss charges you with the wheel spinning.
Signalling: The boss lifts the wheel behind him and spins it.
Evade: Jump over the boss.

Attack 2
Move: The boss tosses the wheel at you diagonally.
Signalling: The boss jumps into the air.
Evade: Slide towards the boss.

Attack 3
Move: Following A2, the boss recalls the wheel which returns to him on the ground, spinning.
Signalling: The boss raises his hand.
Evade: Jump over the wheel.

Strategy
The first boss shouldn't be too difficult for you, regardless of which weapon you have chosen at the beginning. The opportunity to hit the boss comes after you evade A1 or between A2 and A3. Don't attack too many times after A2, or you can get stuck in an animation and A3 will hit you.

Location: Sacred Entombments
Healthpoints estimate: 2000 (not counting the healing)
Tears of Atonement: 2,000
Marks of Martyrdom: 1

Attacks
Attack 1
Move: The boss sweeps his bat low
Signalling: The boss pulls his hand behind him
Evade: Run away from the boss.

Attack 2
Move: The boss smashes his bat at you.
Signalling: The boss pulls his hand behind him, then his eye flashes red.
Evade: Run far away from the boss, as this attack has a far reach.

Attack 3
Move: The boss charges you, then gets stuck in the wall.
Signalling: The boss leans back and glass shards appear on his body.
Evade: Use the wall panels to cling to a wall, then jump over the boss. You need to time it correctly as The Penitent One slides down after a while.

Attack 4
Move: The boss turns around and sweeps his bat (scythe - ?) at you.
Signalling: The boss gets unstuck from the wall.
Evade: Run away from the boss.

Attack 5
Move: The boss sends two (three) purple orbs, alternating between high and low orbs
Signalling: The boss puts a hand in front of him with crossed fingers, and purple light appears in it.
Evade: Jump over the low orbs, and duck or slide under the high orbs.

Attack 6
Move: The boss makes two (three) exploding bubbles appear.
Signalling: The boss puts both hands in front of him with crossed fingers, and the bubbles appear on the ground.
Evade: Run away from the bubbles. Touching them before they explode does not hurt.

Attack 7
Move: Stones drop from the ceiling.
Signalling: The boss climbs up from the ground starts chomping on his bat, or gets stuck on the wall.
Evade: Avoid the stones by running to the side.

Attack 8
Move: The boss sends waves of glass shards on the ground.
Signalling: The boss puts both hands in front of him with crossed fingers.
Evade: Jump over the waves.

Attack 9
Move: The boss throws glass shards at you.
Signalling: The boss raises his left hand behind him.
Evade: Run far away from the boss quickly, as this attack is quite fast though the damage area does not seem that big.

Strategy
In this fight, you can request Yerma's help. She attacks the boss when he is stuck in the wall for quite big chunk of his health.
This boss has two phases. In the first one, he performs A1-A6, and during A5 and A6 he sends the lower number of magical attacks at you. The A2 seems like it can be parried if you have weapon which allows it, but I did not dare to try it yet. After you deplete half of his health bar, he climbs out of the ashes to his full height, and during the transition he also eats the body on his bat which heals him. After that, he does not perform A1 or A2, A4 is changed, and A3 is slightly different, but despite the boss appearing taller it is avoided the same way. During the second phase the boss performs A5-A9, and during A5 and A6 he sends the higher number of magical attacks at you.
The best opportunity to attack is obviously when the boss is stuck in the wall after A3, though during the second phase watch out for the falling stones. You are certain to have the Veredicto for this boss so make use of its damage-dealing capabilities.

Notes
When the boss heals himself, the animation reminds me of the painting "Saturn Devouring His Son" from Francisco Goya.

Location: Palace of the Embroideries
Healthpoints estimate: 1300
Tears of Atonement: 2,000
Marks of Martyrdom: 1

Attacks
Attack 1
Move: The boss stabs across the whole arena.
Signalling: The boss moves to one side, then raises her rapier and lightning appears around it.
Evade: Jump as the boss disappears.

Attack 2
Move: The boss stabs at you.
Signalling: The boss raises her rapier.
Evade: Run away from the boss or parry.

Attack 3
Move: The boss slashes around her.
Signalling: The boss disappears, then a circle of blue flashes appears on you. Sometimes the boss jumps twice to a thread high above the arena before disappearing.
Evade: Run from the flashes which do not hurt you.

Attack 4
Move: The boss spins and throws three (five/seven) bolts at you diagonally.
Signalling: The boss jumps up, then stands on a thread.
Evade: Run from the bolts to put yourself in a gap between them as they separate.

Attack 5
Move: The boss does an upwards slashing move.
Signalling: The boss lowers her rapier behing her.
Evade: Run or slide away.

Attack 6
Move: The boss slashes around her in three (five) places in a row.
Signalling: The boss disappears.
Evade: The damage areas alternate between high and low ones, so crouch under one which is higher.

Attack 7
Move: The boss summons two golden copies which both shoot three, three and five bolts at you.
Signalling: The boss moves to the center of the arena and the copies appear.
Evade: Slide through the bolts.

Strategy
Orospina is the "small and fast" boss archetype. She tends to prevent you from dealing damage to her during the fight a lot, either by jumping high above the arena, or disappearing completely. Good opportunity to hit back is after any of the ground attack (all except A5 and A7). You also need to be careful not to fall in the pits on the sides of the arena. When you deplete two thirds of Orospina's health, she performs A7 as a sort of transition to a second phase, where she becomes a bit faster (this is also reflected in the music which increases in tempo). She also starts to perform the stronger version of A6, though not exclusively.

Location: Crown of Towers
Healthpoints estimate: 800-900 (Lesmes), 500 (Infanta)
Tears of Atonement: 2,000
Marks of Martyrdom: 1

Attacks
Attack 1
Move: Lesmes smashes the coffin into the ground, creating large pillars of fire.
Signalling: The boss lifts the coffin and fiery holes appear on the ground
Evade: Stand between the holes.

Attack 2
Move: Lesmes floats around and has fireballs circle around him, then recalls the fireballs and finishes with A1.
Signalling: The boss rises into the air, then transforms with an explosion.
Evade: The fireballs move a bit tricky so try to avoid them as the boss summons them and start to rotate them. Don't stand in the fireball circle when the boss is about to recall the fireballs. Avoid A1 as usual.

Attack 3
Move: Lesmes smashes the coffin into you.
Signalling: The boss raises the coffin behind him, then dashes towards you with orange shadow behind.
Evade: Jump over or slide through the boss.

Attack 4
Move: Lesmes smashes the coffin into the ground three times, creating a fire pillar.
Signalling: The boss rises into the air, while holding the coffin.
Evade: The boss follows you so keep moving. The pillars are quite large and you can find yourself boxed in, but you can just about slide through one of the pillars, so do that if you appear to be blocked. Alternatively, move only a bit to the side when the boss starts to drop as he will make a pillar just next to the previous one.

Attack 5
Move: Infanta sends the miasma spikes at you, one by one.
Signalling: The boss raises a hand which glows green.
Evade: The spikes attack the place where you were standing a moment ago so keep jumping and ducking to avoid them.

Attack 6
Move: Infanta sends the miasma spikes at you, one by one.
Signalling: The boss raises a hand which glows green.
Evade: The spikes attack the place where you were standing a moment ago so keep jumping and ducking to avoid them.

Attack 7
Move: Infanta sends the remaining miasma spikes at you as a volley.
Signalling: The boss raises a hand.
Evade: Run from the spikes.

Attack 8
Move: Infanta charges across the arena with the spikes rotating around her.
Signalling: The boss lands, leans forward and the spikes start to rotate.
Evade: Jump over the boss.

Strategy
Lesmes' hitpoints are hard to guess as the fire effect of his attacks covers up the damage numbers. There is not much to his fight, except A2's fireball movement (as noted above).
Once you deplete Lesmes' healthbar, the fight continues as Infanta climbs out of the coffin and you need to deal with her. She attacks using six miasma spikes which she summons around her. Her attacks are A5-A8. Sometimes, she performs the attacks while floating in the air.
Once you defeat Infanta as well, Lesmes returns and you have to face both bosses at the same time. Sometimes, they team up against you and perform A3 and A8 at the same time. Otherwise, they randomly perform their attacks, same as during their separate fights. Start by focusing on defeating Infanta, as she has lower hitpoints. Once she is down, deal with Lesmes as before.

Notes
"This boss wasn't so bad... Oh."

Location: Under Her Sacred Grounds - you can reach it through Mother of Mothers before defeating any of the guardians, you just need to obtain all three weapons
Healthpoints estimate: 2500
Tears of Atonement: 2,500
Marks of Martyrdom: 2

Attacks
Attack 1
Move: The boss charges towards you spinning (five times).
Signalling: The boss puts his sword hand under his other hand.
Evade: Slide through the boss when he is close.

Attack 2
Move: The boss swings his sword downwards at you (and creates a damaging wave).
Signalling: The boss sharpens the sword, then puts it above his head.
Evade: Run away from the boss. Slide through the wave.

Attack 3
Move: The boss jumps forwards or backwards spinning, then on landing creates damaging explosions on the floor.
Signalling: The boss holds his sword above his head with both hands. The jump direction depends on his and your position.
Evade: Stay away from the boss as he jumps, then jump over the explosions.

Attack 4
Move: The boss spins and throws five knives at you which ricochet three times off the walls and ground in a random pattern.
Signalling: The boss starts to spin slowly, then flashes appear where the knives will strike
Evade: Try to guess the path of the knives based on the flashes, then avoid standing in the path.

Attack 5
Move: The boss drops random sharp weapons from the ceiling
Signalling: The boss breaks his sword when doing A2, then jumps into the background.
Evade: Run to the sides as the weapons fall; The pattern is random but there aren't that many.

Attack 6
Move: The boss drops from the ceiling and creates damaging explosions on the floor.
Signalling: The boss jumps from the background.
Evade: The boss will fall where you are so keep running to one side, then jump if needed to avoid the explosions.

Attack 7
Move: The boss drops from the ceiling and creates a damaging wave.
Signalling: The boss holds his sword above his head with both hands, then jumps directly up.
Evade: The boss falls on one side of the arena, so stand in the middle. Slide throught the wave.

Attack 8
Move: The boss throws knives at you.
Signalling: The boss spins and floats in the middle.
Evade: Stand to the side of the arena.

Strategy
In this fight, you can request Yerma's help. She attacks the boss when he is close to a wall.
This boss has three phases corresponding to the thirds of his health bar. In the first one, he performs A1-A4, where A1 and A2 are the weaker versions. After you deplete a third of his life, he performs A5 followed by A6, which serves as a transition to the second phase where he starts to perform the stronger version of A2. Upon depleting two thirds he performs A5 and A6 again, then in the last phase he starts to perform the stronger version of A1 (but sometimes makes only one spinning charge at you, so be on your toes for repeated charges) and also adds A7. The boss is not so fast so after each of his attacks you have a chance to damage him.

Notes
This is probably my favourite boss fight. I like the whistling sound the boss makes while sharpening his sword, and also the detail where the sword first loses its rust, then becomes gleaming gold and sharp as the fight progresses.
A8 was added in the Mea Culpa/True Torment update, I believe it was not there before.

Location: Basilica of the Absent Faces
Healthpoints estimate: 1700
Tears of Atonement: 3,000
Marks of Martyrdom: 3

Attacks
Attack 1
Move: The boss summons eight gray orbs around you which chase you, dealing damage on contact.
Signalling: Gray sparks appear on the boss' chest; Orbs start to appear around you clockwise.
Evade: Dash through the orbs, then run away (the orbs are rather slow)

Attack 2
Move: The boss shoots a short lightning bolt at you, three times.
Signalling: The boss gains a blue aftershadow and dashes across the arena, following you, then raises one hand.
Evade: Jump away when the boss stops moving.

Attack 3
Move: The boss summons coffins which move across the screen vertically.
Signalling: The boss disappears, and the coffins start to appear at the bottom of the screen.
Evade: The coffins start to move in the reverse order in regard to when they appear. Unfortunately I don't know the exact method to evade it as the attack covers basically the whole screen. Dodging through the already moving coffin to avoid the next one seems like the best bet, but it did not work for me very well.

Attack 4
Move: The boss summons fewer coffins which move across the screen vertically in three waves
Signalling: Gray sparks appear on the boss' chest; Coffins start to appear on the bottom and the top of the screen
Evade: Stand in a gap between the coffins. Move between waves

Attack 5
Move: The boss destroys one platform.
Signalling: The boss moves to the chain holding the platform and strikes it three times with A2.
Evade: Jump to the other platform.

Attack 6
Move: The boss summons three coffins which move across the screen horizontally
Signalling: The boss moves to the edge of the arena and raises a hand.
Evade: Jump over the coffins. Note that similarly to the area's special feature, the coffins which go off one side of the screen come back from the other side, so you have to dodge each coffin twice.

Strategy
The fight takes place on floating platforms. Falling off means certain death, so watch out especially for A5 which can easily kill you if you mistake it for A2. The boss performs it after you remove about half of her hitpoints. After that, your space is even more limited. Use Veredicto's reach to deal with the boss which floats around the arena.

Notes
Unfortunately the fight was reworked a bit in the Mea Culpa/True Torment update (I wrote the core of this guide before that). However, the attacks are mostly the same.

Location: Sea of Ink/Sunken Cathedral
Healthpoints estimate: 700 (first encounter), 2300 (actual fight)
Tears of Atonement: 3,500
Marks of Martyrdom: 3

Attacks
Attack 1
Move: The boss performs an uppercut with his halberd (sending a wave of water in each direction).
Signalling: The boss steps towards you and prepares his halberd.
Evade: Jump over the boss.

Attack 2
Move: The boss jabs his halberd at you (sending a wave of water).
Signalling: The boss raises the halberd behind his back.
Evade: Run or slide away from the boss.

Attack 3
Move: The boss slashes his halberd at you while turning around (sending a wave of water).
Signalling: The boss, who has his back at you, starts to turn around.
Evade: Run or slide away from the boss.

Attack 4
Move: The boss drops from the ceiling, then creates two water waves, one to each side.
Signalling: The boss jumps up, then a circle of water appears of the ground beneath you.
Evade: The circle of water indicates where the boss will fall, so run from it. Jump over the wave.

Attack 5
Move: The boss summons waves of pikes from the ground.
Signalling: The boss kneels, then raises his halberd.
Evade: Jump over the waves.

Attack 6
Move: The boss summons two (three) pikemen who appear in a flash of lightning.
Signalling: The boss kneels, then raises his halberd with a flash of lightning going to the sky.
Evade: The pikemen appear where you stand, so run around to avoid the lightning and the contact damage from the pikemen. Then beat the pikemen.

Attack 7
Move: The boss puts a shield on the summoned pikemen.
Signalling: The boss kneels, then raises his halberd and a ghostly flag appears on it.
Evade: Damage the boss.

Strategy
You encounter the boss first when going through the Sea of Ink. This fight is shorter and you are not presented with the boss' healthbar. In that fight, the boss performs A1-A5 only (A1-A3) are the weaker versions.
The final fight happens after you get through the Sunken Cathedral. The boss starts performing the stronger versions of A1-A3 now and also can summon pikemen enemies, so deal with them first (unless the boss performs A7, in which case go for the boss). Otherwise there is not much to it, just note that most attacks have sort of "after-attack" (usually a wave of water on the ground).

Notes
I hate when bosses summon smaller enemies to help them.

Location: Severed Tower
Healthpoints estimate: 2900 (plus healing)
Tears of Atonement: 5,000
Marks of Martyrdom: 4

Attacks
Attack 1
Move: The boss creates three (one in the middle, then two on the sides) miasma columns that rise from the ground, then explode and send outward miasma waves across the ground.
Signalling: The old man face shouts and miasma whirlpools appear on the ground around you.
Evade: Crouch to avoid the first column. After the two columns disappear, jump to avoid the miasma waves,

Attack 2
Move: The boss creates several fire orbs which shoot flames to the ground.
Signalling: The old man face shouts and fire orbs appear in the air.
Evade: Stand between the orbs.

Attack 3
Move: The boss shoots a short fire orb volley to four directions, swapping faces.
Signalling: The boss' face gets covered in flames, then explodes.
Evade: Run to the side so you can avoid the orbs more easily.

Attack 4
Move: The boss transforms into a pillar of fire, shooting fire waves on the ground.
Signalling: The nun face is lowered.
Evade: Dash through the pillar, jump over the waves.

Attack 5
Move: The boss shoots flames from the middle two small side faces.
Signalling: The two faces' eyes glow.
Evade: Stand under the face.

Attack 6
Move: The boss heals part of its health.
Signalling: Halo of fire appears behind the nun face and the boss shakes
Evade: No need, this move does not pose danger to you.

Attack 7
Move: The boss shoots long volley of fire orbs in two directions from its mouth.
Signalling: The masked face opens its mouth.
Evade: Stand under the face, the orbs should not go there.

Attack 8
Move: The boss shoots long volley of fire orbs in four directions from its mouth (followed by two more volleys from random places in the air.).
Signalling: The masked face opens its mouth.
Evade: Run away from the boss to gain manoeuvering space. The harder version of the attack is quite bullet hell-like, so try to navigate the gaps between the moving orbs.

Attack 9
Move: The boss covers the floor in flames.
Signalling: The boss transforms into a melted face; Cherub handholds appear in the air.
Evade: Jump on the handholds.

Attack 10
Move: The boss sends a pillar of flame across the arena.
Signalling: None immediate, except it occurs after the arena transformation during A8.
Evade: Dash through the pillar.

Strategy
The weak point of the boss is the face. The boss swaps the faces, and each of them performs different set of attacks. There are not thousand of them though as the name would suggest, only four: old man (initial, does A1 and A2), nun (does A4-A6), iron mask (does A7 and A8, randomly with harder version) and melted face (final, does A9 and A10). The attacks are mostly fire-based.
Good time to deal damage is during A5, and also A6. Usually after any attack the boss is open to damage as well. After you reduce most of the boss' healthbar, it transforms the arena with A9. Now you cannot stand on the ground, you have to jump between the handholds and avoid A10, then use your air movement abilities to reach the face and hit it (I'd say the fast-hitting weapons are better at this).

Location: Two Moons
Healthpoints estimate: 2900-3000
Tears of Atonement: 7,500
Marks of Martyrdom: 4

Attacks
Attack 1
Move: The boss floats around the arena shooting lightning orbs, three per volley.
Signalling: The boss starts to float from the edge of the arena.
Evade: Avoid the orbs

Attack 2
Move: The boss sends three lightning columns across the arena, twice.
Signalling: The boss moves to the edge of the arena and gets restored to the original state (face instead of skull, full body).
Evade: Slide through the columns.

Attack 3
Move: The boss shoots a wide lightning beam downward and moves across the arena (and back).
Signalling: The boss moves to the edge of the arena and starts to glow
Evade: Thin platforms appear in the air, so hide below them. Watch out as the platforms that appear are narrower and narrower as the fight progresses.

Attack 4
Move: The boss shoots a volley of orbs
Signalling: The boss moves to the edge of the arena and gets restored to the original state (face instead of skull, full body).
Evade: Jump over the low orbs, duck under the high orbs. For the volley of two rows, double jump and dash.

Attack 5
Move: The boss shoots a bullet-hell-like volley of orbs
Signalling: The boss moves to the edge of the arena and gets restored to the original state (face instead of skull, full body). Boss' heart glows blue.
Evade: Jump precisely through the orbs.

Strategy
You can enlist Yerma's help for the last time here. She attacks the boss when it gets to the edge of the screen.
Beads providing lightning resistance are certainly a good idea here. Otherwise there is not much to it, jump and hit the boss when it is floating around during A1.

Notes
I got the "Defeat a boss without being hit" achievement in this fight, so perhaps it is a good opportunity to get it as the fight is not that hard.
A4 and A5 were added in the Mea Culpa/True Torment update. I don't have them recorded and have only seen A5 once, so the info might be incomplete. My apologies.

Location: Crimson Rains
Healthpoints estimate: 1700
Tears of Atonement: -
Marks of Martyrdom: -

Attacks
Attack 1
Move: The boss shoots a lightning waves towards you.
Signalling: The boss appears at the edge of the arena and raises the hand with the nail.
Evade: Slide/dash through the waves.

Attack 2
Move: The boss throws a large fireball to the ground, which explodes and creates fire columns around it.
Signalling: The boss appears above you and raises a hand.
Evade: Run to the side to avoid the fireball, double jump over the columns.

Attack 3
Move: The boss shoots several red orbs and one green orbs.
Signalling: The boss appears about above the edge of the healthbar (either side).
Evade: Slide/dash through the orbs.

Attack 4
Move: The boss teleports and shoots two red orbs, three times.
Signalling: The boss appears and raises his hand
Evade: Slide/dash through the orbs.

Strategy
After each attack (maybe with the exception of A2), the boss floats in the air doing nothing, so use this opportunity to deal damage or heal.
Notes
Easy fight, but it is just the first phase, and you have to do it every time you want to attempt the second, so it is better to do it with as little damage/healing as possible.

Location: Crimson Rains
Healthpoints estimate: 3000
Tears of Atonement: 10,000
Marks of Martyrdom: 5

Attacks
Attack 1
Move: The boss performs a three attack combo: first, he dashes toward you and finishes the dash with an uppercut. Then, he teleports (usually behind you, but sometimes in front, usually if you are close to the arena's edge) and does a low cut. Last, he creates two shockwaves on the ground, one to each direction.
Signalling: The boss puts his sword behind him, holding it diagonally upwards.
Evade: Double jump or keep distance from the first attack (it has limited range). It can also be blocked. Ideally, parry the second attack as doing that cancels the third attack and deals decent damage. If you do not do that, jump or slide/dash through the third attack. Watch out as the shockwaves become bigger right before disappearing and this can land a hit on you.

Attack 2
Move: The boss smashes into the ground, causing red columns to raise.
Signalling: The boss leaps into the air towards you.
Evade: Slide/dash towards the boss.

Attack 3
Move: The boss sends red shockwaves from his sword, usually twice in a row.
Signalling: The boss teleports and raises his sword in front of him. Red lightning appears above him.
Evade: Jump over the shockwaves.

Attack 4
Move: The boss performs a three attack combo: first, he dashes toward you and finishes the dash with an uppercut and leaps into the air. Then, he does an area-of-effect slash around him. Last, he smashes into the ground and creates red columns, similarly to A2.
Signalling: The boss puts his sword behind him, holding it diagonally upwards, then putting it on the ground. During the dash, small lightning bolts appear on the ground.
Evade: Keep distance from the first attack (it has limited range). It can also be blocked. Slide under the second attack, then slide/dash from the ground smash and jump over the columns.

Attack 5
Move: The boss does a large area-of-effect slash around him.
Signalling: The boss teleports and appears high in the air, usually in the middle of the arena.
Evade: Try to run from the area of effect, but the time is rather short, so it is better to slide/dash as the boss attacks

Attack 6
Move: The boss summons the previous bosses: Orospina, Lesmes, Infanta, Benedicta and Odon for one attack each. Orospina slashes across the arena, Afilaor drops from the ceiling and creates a large fire column, Infanta shoots a volley of miasma spikes, Benedicta drops the coffins across the arena (vertically) and Odon drops from the ceiling and performs a dashing uppercut. The boss finishes this with A5.
Signalling: The boss finishes one of his combos with creating a large red and black column, similarly to what happened in the beginning of the fight, and the screen darkens.
Evade: Avoid the red and black column as it can damage you. Jump over Orospina, run from the fire column, slide/dash through the green spikes, slide/dash as the coffins drop to avoid them, jump over Odon as he charges. Avoid the A5 as usual.


Strategy
Well, this is it. Probably the hardest fight in the game. The strategy that worked quite well for me was using Ruego al Alba to block the first hit of A1, then parry the second hit of the same attack. Combined with items to boost RaA's damage and prolong the block window of opportunity, the boss did not stand much chance. There are two caveats for this: one, the boss sometimes teleports behind you for the second hit, so you have to turn around quickly and parry. Second, resist activating the blood pact mode for the weapon, as that takes away your ability to parry which at least for me was crucial. The parry deals decent damage though anyway, and on top of that, cancels the third hit. The boss is quite fast, but if you learn the attacks as described above and utilize your movement abilities, you should be able to avoid most damage. After A2, there is a good opportunity to deal damage or heal, if you lost some health. The difference between signalling for A4 and for A1 is quite small, but if you block the first hit, you can adapt to the second hit based on whether the boss disappeared (A1) or launched into the air (A4).
Notes
In the early days after game's release, it was possible to cheese this fight by using the time stop prayer and recovering enough mana to keep the time stopped for the full duration of the fight, which prevented the boss from fighting back. The prayer was altered in a later patch to have a shorter duration.

Location: Crimson Rains
Healthpoints estimate: 4000 (Boss), 100 (flowers)
Tears of Atonement: 0
Marks of Martyrdom: 0

Attacks
Attack 1
Move: The boss sends two blue lightning columns across the arena, one from each side.
Signalling: The heart in the middle flashes blue.
Evade: Dash through the first, then the second column.

Attack 2
Move: The boss sends two blue lightning columns from the middle of the arena to the edges.
Signalling: The heart in the middle flashes blue.
Evade: Dash through the column threatening to hit you.

Attack 3
Move: The boss creates a row of fire explosions at the bottom of the arena.
Signalling: A row of fire orbs appears on the ground.
Evade: Double jump over the explosions.

Attack 4
Move: The boss covers the ground with blue flames.
Signalling: Four platforms appear in the air.
Evade: The flames go from the sides to the middle, so try not to stand at the edges. Jump on a platform. The flames stay until you progress a bit with the fight.

Attack 5
Move: The boss summons four flowers which repeatedly shoot white orbs at you (three per volley).
Signalling: The heart is covered with a golden shield and becomes invulnerable.
Evade: Dash through or destroy the orbs and destroy the flowers. Try to do it from edge to edge as the flowers respawn after a while.

Attack 6
Move: The boss drops debris at you from one edge of the arena to the other, four times in a row.
Signalling: Columns of light appear across the arena.
Evade: Stand between the columns to avoid the debris

Strategy
The weak point of the boss you need to hit is the heart in the middle. The boss starts with A1, A2 and A3. After losing a bit of health, the boss performs A4 and the arena transforms a bit. Then they still perform the same three attacks, but you need to stay on the four platforms and avoid the flames below, so it is slightly harder. After losing some more health, the boss performs A5 so destroy the flowers quickly (because you cannot damage the heart as long as at least one is alive). Then, the flames and platforms should disappear and the boss should perform A6. After that, it is back to the original three attacks and possibly even the A4 can appear again. Use the downtime after each attack to hammer at the heart (or heal if you need to) and it should go down with no problems.

Notes
And that's it, you have beaten the game!

• Add remaining boss screenshots
  
• Add DLC bosses and minibosses
  
• Add elemental type of each attack (uncertain)
  
• Add weaknesses of the bosses (uncertain)

1.0 (03/11/2024): Initial version
1.1 (17/11/2024): Added most screenshots for most base game bosses; Added ToA/MoM rewards;Updated sections of bosses which were changed in the Mea Culpa/True Torment update