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Research Tier IV->Max lvl->Max boost
Damage II->5->250%
Kill Exp (%)->5->25%
Gold Bonus Per Wave->10->10
Gold Drop (%)->5->50%
Mob Gold->5->125%
Tank Gold->5->250%
Tech Points->3->+9
That double damage is on each damage upgrade seperate.
After reaching wave 3500 the guide here is good for further prioritizing.
But dont use it before wave 3500, it only make your game advancing extremely slow.
Explanation:
It is mandatory to get to wave 3500 as the first goal.
Nothing give you more upgrades to everything then the artifacts and then reroll all on wave 2500 and later 3500.
Only the damage upgrades give you this without grinding month of playing.
Sure you can upgrade like the above guide and upgrade your towers to a high level to get to wave 3500. Will need a few month.
Or you invest in the damage upgrades a little first and you have wave 3500 in weeks.
And then you can upgrade your towers in days to the same outcome with the extremly boost of XP you get of the artifacts.
Every different damage upgrade doubles your damage if you double the outcome of the upgrade.
e.g.
damage upgrade to the first 100% more damage = double damage
The same to 200% = double damage again.
The same to 400% = double damage again.