Out of all the female leads in the series, the goddesses are the ones I sympathize with most. They never get a happy ending despite all the sacrifices. At the end of Origin, they seal the Black Pearl only to be awakened and seal it again. To my knowledge, Ys 2 is where their story truly ends. They are just sealed along with the Black Pearl.

Don't get me wrong, I feel bad for the other Winged Ones too but some of them deserved it and one could argue that some of them were liberated from their curse. These 2 just wanted to live with their friends. Not to mention their relationship with Adol and Toal respectively.